== Vernacular names ==
* *ssp. ''longan'' var. ''longan'' : *longan (En). *Longanier, oeil de dragon (Fr). *Indonesia, Malaysia: lengkeng
*Burma: kyet mouk
*Cambodia: mien
*Vietnam: nhan.
*ssp. ''longan'' var. ''obtusus'' : *Thailand: lamyai khruer, lamyai tao.
*ssp. ''malesianus'' var. ''malesianus'' : *Malaysia: mata kucing (Peninsular Malaysia and Sabah), isau, sau, kakus (Sarawak)
*Indonesia: buku, ihau (Kalimantan), medaru (Sumatra).
== Properties ==
The edible portion of export quality fruit of 3 cultivars ranges from 67 to 78% of the whole fruit. Composition of longan per 100 g edible portion is: water 72.4 g, protein 1.0 g, fat 0.5 g, carbohydrates 25.2 g, fibre 0.4 g, ash 0.5 g, Ca 2 mg, P 6 mg, Fe 0.3 mg, vitamin A 28 IU, vitamin B<sub>1</sub>0.04 mg, vitamin B<sub>2</sub>0.07 mg, niacin 0.6 mg and vitamin C 8 mg. The energy value averages 458 kJ/100 g. The sugar content is very high. The composition of "mata kucing" fruit is not very different, but carbohydrates - and energy values - are much lower, whereas much higher figures are given for mineral content.
== Description ==
*Tree, up to 40 m tall and 1 m trunk diameter, sometimes buttressed, exceptionally a scandent shrub; branches terete with 5 faint grooves, sometimes warty lenticellate, rather densely ferruginous tomentose. *Leaves 2-4(-6)-jugate, axial parts mostly densely hairy; petiole 1-20 cm, petiolules 0.5-35 mm long; leaflets elliptical, 3-45 cm × 1.5-20 cm, 1-5 times longer than wide, chartaceous to coriaceous, above often tomentose in basal part of midrib, beneath thinly tufted-tomentose mainly on midrib and nerves. *Inflorescences usually terminal, 8-40 cm long, densely tufted-tomentose; cymules (1-)3-5-flowered; pedicels 1-4 mm; bracts patent, 1.5-5 mm long; flowers yellow-brown; calyx lobes 2-5 mm × 1-3 mm; petals 5, 1.5-6 mm × 0.6-2 mm, densely woolly to glabrous; stamens (6-)8(-10), filament 1-6 mm. *Fruit drupaceous, 1-3 cm in diameter, lobe(s) broad-ellipsoid to globular, smooth to warty or sometimes up to 1 cm aculeate, sometimes granular, glabrescent, yellow-brown. *Seed globular with shining blackish-brown testa; seed enveloped by a thin fleshy, translucent white arilloid.
== Growth and development ==
Longan trees grown from air layers come into bearing during the third or fourth year and yields tend to increase with tree size over a very long trajectory. Flowering within a panicle is a sequence of opening of staminate (pistil non-functional), pistillate (stamens non-functional), hermaphrodite and finally staminate flowers again. Male and female phases of flowering overlap 4-6 weeks depending on cultivars. Pollination, by insects, is most effective between 8.00 a.m. and 2.00 p.m. In one study fruit set per panicle improved greatly with bloom rating for the tree, leading to a sharp progression in yield per tree (and an obvious risk of biennial bearing). The period from bloom to harvest is 5-7 months, depending on cultivar and climate. Longan in Thailand flowers just before or after the temperature rise at the end of the cold, dry season. Most fruit is harvested in August and September.
In the panicles of "mata kucing" male and hermaphrodite flowers occur. Flowering lasts only 1-2 weeks, the period varying from late June to mid-October in Sarawak. Pollination is mainly by insects such as ants, flies and honey bees ( ''Apis cerana'' , ''A. florea'' and also ''A. dorsata'' ). Flowering is often prolific but fruit set is quite low and 2 weeks after flowering, when fruitlets measure 1 cm, many are shed. The fruit ripens about 4 months after bloom.
== Other botanical information ==
The two subspecies and five varieties of ''D. longan'' , listed above, are distinguished mainly by differences in the leaflets.
Within ssp. ''malesianus'' , var. ''malesianus'' shows the greatest variation in Borneo. The fruits are globular to slightly oblong and smooth to warty. In Peninsular Malaysia, the most common form of this taxon is the one with globose smooth fruits which turn brown when ripe. This is the true "mata kucing" and has usually been identified as ''Euphoria malaiensis'' . It has a very thin arilloid and is hardly worth eating. This form also exists in Borneo and Sumatra. The more superior forms are found in Sarawak, all with densely thick warty fruits and thicker arilloids. These forms can be roughly grouped into three types based on the fruit characteristics: the "isau" with fruits which are globular and remain green when ripe, the "sau" with fruits which are slightly oblong and also remain green when ripe, and the "kakus" with globular fruits which turn brown when ripe. The leaves, flowers and tree forms also differ. The "kakus" is more widespread in Sarawak, while the "isau" and "sau" are mainly confined to the river banks of the Rajang river and to the Bareo valley.
Var. ''echinatus'' differs from var. ''malesianus'' in that the fruits have rather long spines resembling the rambutan ( ''Nephelium lappaceum'' L.). This variety is found in Sabah where the "kakus" also exists.
Three edible longan types are distinguished in Thailand, which presumably all belong to ssp. ''longan'' . The first one is a large forest tree with small fruits and a very thin aril, possibly of interest for breeding purposes. The second one is the common longan ("lamyai kraduk"), growing in the northern part of the country as an erect tree, producing small fruits with large seeds and is recommended as a rootstock for commercial cultivars. The third type is formed by the commercial cultivars ("lamyai kraloke") which produce large fruits and small seeds.
Longan thrives on rich sandy loams, it does well on oolitic limestone; moderately acid sandy soils are more marginal and on organic muck soils flowering is deficient, probably because shoot growth continues for too long. In northern Thailand longan orchards are often situated on the lighter soils along former river courses, a ribbon of trees winding between the sawahs. The roots grow down 2-4 m to the water table.
The "mata kucing" thrives in the humid tropical lowlands near sea level, within about 10°from 10° from the equator. The trees occur mainly in the substage or understorey in primary or sometimes secondary forests. Rainfall ranges from 2500 mm to more than 4000 mm per year associated with a mean air temperature of 25-30°C and a relative humidity of 65-95%. In Sarawak, the trees grow on alluvial soil, often on river banks. In other areas the trees grow on a wider range of soil types. A pH range of 4.5-6.5 is common in this region.
== Propagation and planting ==
Seeds are too short-lived for germplasm collection. Thailand has large tree collections of longan in Chiang Mai and Lamphun. The Thai cultivars differ in shoot, flower and fruit characters from the Chinese cultivars, but on the whole, genetic diversity appears to be narrow. There are several cultivar collections in Australia, the largest being in Kamerunga Horticultural Research Station near Cairns, Queensland.
The University of Agriculture Malaysia with its branch campus at Bintulu, Sarawak, is now the largest collector of germplasm of ''D. longan'' ssp. ''malesianus'' . The great diversity in Sarawak offers a great opportunity to select superior material. Explorations in remote areas have been regularly made to identify trees with good quality fruit - thick flesh, fruit in consolidated panicles - and to collect budwood.
== Breeding ==
If trees bore regularly, growth would be moderated and it would be easier to prune to keep trees a manageable size. Small trees, coupled with closer spacing and regular yields would allow production to be intensified.
The superior races of the spp. ''malesianus'' , in particular the var. ''malesianus'' in Sarawak and other parts of Borneo, may offer an attractive alternative to longan for the humid tropical lowlands.
== Literature ==
* Anonymous, 1987. Lychees and longan. Union Offset, Bangkok. pp. 44-71. (Thai).
* Tongdee, S.S., 1977. Study on the characteristics of longans during storage. Kasikorn 50(2): 95-97. (Thai).
* Verheij, E.W.M. & Koopmans, A., 1984. Flowering and fruiting of longan (Euphoria longana Lam.) in East Java in 1983. Agrivita 7(1): 14-19.
* van Welzen, P.C., Lamb, A. & Wong, W.W.W., 1988. Edible Sapindaceae in Sabah. Nature Malaysiana 13: 10-25. | '''10''' | * Wong, K.C., Ibrahim Yusof, Pearce, K.G. & Alau Tayan, D., 1988. Isau - A potential tropical longan (Dimocarpus longan) of Sarawak. Proceedings of the Third National Biology Symposium, Subang Jaya (in print).
== Authors ==