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Brachystegia spiciformis (PROTA)

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<big>''[[Brachystegia spiciformis]]'' Benth.</big>
 
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{{DISPLAYTITLE:''Brachystegia spiciformis'' (PROTA)}}
 
:Protologue: Trans. Linn. Soc. London 25: 312 (1866).
In traditional charcoal making in Tanzania, the average yield of marketable charcoal from ''Brachystegia spiciformis'' wood with 40–50% moisture content was about 90 kg per m³. The foliage is of intermediate value as forage. The rumen degradability is 0.4–0.5. Young leaves contain per 100 g dry matter approximately: digestible dry matter 52 g, crude protein 18 g and energy 760 kJ; for old leaves: digestible dry matter 60 g, crude protein 11 g and energy 870 kJ.
== Botany Description ==
Deciduous or semi-evergreen, small to fairly large tree up to 35(–40) m tall; bole branchless for up to 15 m, straight and cylindrical or poorly formed, up to 120 cm in diameter, without buttresses but sometimes fluted at base; bark surface smooth, grey to greyish brown, often flaking off in thick scales, in older trees becoming shallowly fissured, inner bark reddish; crown rounded but becoming flattened in older trees, with spreading branches; twigs glabrous or slightly hairy. Leaves alternate, paripinnately compound with 2–8 pairs of leaflets; stipules 1–4 cm long, usually caducous; petiole 1–5 cm long, thickened at base, rachis (3–)5–15(–18) cm long, grooved and slightly winged; leaflets opposite, ovate to elliptical, oblong or lanceolate, 1.5–20 cm × 0.5–8 cm, basal pair of leaflets smallest, obliquely rounded, truncate or cordate at base, acute to rounded or notched at apex, glabrous or slightly hairy, pinnately veined but with 3–5 basal veins. Inflorescence a terminal or axillary panicle up to 8 cm long, glabrous to short-hairy. Flowers bisexual, nearly regular, small, fragrant, at base with 2 obovate to rounded bracteoles 0.5–1 cm long; pedicel 2–8 mm long; sepals 0–4, slightly unequal, 0.5–3 mm long, nearly glabrous; petals absent; stamens 10, fused at base, 1–2 cm long; ovary superior, ellipsoid, 2–5 mm long, with stipe up to 6 mm long, hairy, style slender, up to 1.5(–2) cm long, coiled. Fruit an oblong to obovoid, flattened pod 8–15(–20) cm × 3–5 cm, at a right angle to the stipe, smooth but slightly wrinkled, slightly winged along one suture, dark brown to purplish brown, dehiscent with 2 woody valves, up to 6-seeded. Seeds oblong to ovoid, 1.5–2.5 cm long, dark brown.
 
== Other botanical information ==
''Brachystegia'' is a taxonomically difficult genus comprising about 30 species, distributed in mainland tropical Africa and South Africa, the majority of species occurring in southern tropical Africa, where they are characteristic of miombo woodland.
''Brachystegia spiciformis'' is extremely variable in leaf and inflorescence characters. It is slow growing, and may become a fairly large tree in wetter savanna and woodland, but remains a small stunted tree in dry areas. The taproot can reach a depth of more than 5 m and laterally the root system can extend over more than 25 m. Flowering and fruiting do not always occur every year and depend on the climatic conditions. In southern Africa, flowering usually occurs in August–November. The flowers are much visited by bees, which probably pollinate them. After pollination, fruit development takes 7–8 months. At the end of the dry season seeds are scattered when dry fruits split open explosively.
 
== Description ==
 
 
 
== Other botanical information ==
 
 
 
== Growth and development ==
 
 
== Ecology ==
''Brachystegia spiciformis'' occurs in coastal and upland deciduous woodland and open forest, up to 2000(–2350) m altitude. It is characteristic of miombo woodland, where it is often dominant in higher rainfall areas. It can be found on hill slopes and river banks, but also on intensely drained and very poor and shallow soils. The annual rainfall in the area of distribution ranges from 500 mm to 1200 mm. ''Brachystegia spiciformis'' is frost and fire tender.
 
== Propagation and planting ==
 
 
== Management ==
* Brummitt, R.K., Chikuni, A.C., Lock, J.M. & Polhill, R.M., 2007. Leguminosae, subfamily Caesalpinioideae. In: Timberlake, J.R., Pope, G.V., Polhill, R.M. & Martins, E.S. (Editors). Flora Zambesiaca. Volume 3, part 2. Royal Botanic Gardens, Kew, Richmond, United Kingdom. 218 pp.
* Coates Palgrave, K., 2002. Trees of southern Africa. 3rd Edition. Struik Publishers, Cape Town, South Africa. 1212 pp.
* Lovett, J.C., Ruffo, C.K., Gereau, R.E. & Taplin, J.R.D., 2007. Field guide to the moist forest trees of Tanzania. [Internet] Centre for Ecology Law and Policy, Environment Department, University of York, York, United Kingdom. http://celp.org.uk/ projects/ tzforeco/. November 2011.
* Mbuya, L.P., Msanga, H.P., Ruffo, C.K., Birnie, A. & Tengnäs, B., 1994. Useful trees and shrubs for Tanzania: identification, propagation and management for agricultural and pastoral communities. Technical Handbook 6. Regional Soil Conservation Unit/SIDA, Nairobi, Kenya. 542 pp.
* Orwa, C., Mutua, A., Kindt, R., Jamnadass, R. & Simons, A., 2009. Agroforestree database: a tree reference and selection guide. Version 4.0. [Internet] World Agroforestry Centre (ICRAF), Nairobi, Kenya. http://www.worldagroforestry.org/ resources/databases/ agroforestree. November 2011.
== Other references ==
* Campbell, B. (Editor), 1996. The miombo in transition: woodlands and welfare in Africa. Centre for International Forestry Research (CIFOR), Bogor, Indonesia. 266 pp.
* Chinuwo, T., Gandiwa, E., Mugabe, P.H., Mpofu, I.D.T. & Timpong-Jones, E., 2010. Effects of previous cultivation on regeneration of Julbernardia globiflora and Brachystegia spiciformis in grazing areas of Mupfurudzi Resettlement Scheme, Zimbabwe. African Journal of Range & Forage Science 27(1): 45–49.
* Dale, I.R. & Greenway, P.J., 1961. Kenya trees and shrubs. Buchanan’s Kenya Estates Limited, Nairobi, Kenya. 654 pp.
* Ernst, W.H.O., 1998. Seed and seedling ecology of Brachystegia spiciformis, a predominant tree component in miombo woodlands in South Central Africa. Forest Ecology and Management 25(3 4): 195–210.
* Grundy, J.M., Campbell, B.M. & Frost, P.G.H., 1994. Spatial pattern, regeneration and growth rates of Brachystegia spiciformis and Julbernardia globiflora. Vegetatio 115: 101–107.
* Herd, A.R.C., 2007. Exploring the socio-economic role of charcoal and the potential for sustainable production in the Chicale Regulado, Mozambique. MSc thesis, University of Edinburgh, United Kingdom. 76 pp.
* Luoga, E.J., Witkowski, E.T.F. & Balkwill, K., 2000. Economics of charcoal production in miombo woodlands of eastern Tanzania: some hidden costs associated with commercialization of the resources. Ecological Economics 35(2): 243–257.
* Nshubemuki, L. & Mbwambo, L., 2007. Trees to promote in the management of miombo woodlands in Tanzania: Species, sizes and qualities. [Internet] MITMIOMBO – Management of Indigenous Tree Species for Ecosystem Restoration and Wood Production in Semi-Arid Miombo Woodlands in Eastern Africa. Proceedings of the First MITMIOMBO Project Workshop held in Morogoro, Tanzania, 6th–12th February 2007. Working Papers of the Finnish Forest Research Institute 50: 57–63. http://www.metla.fi/ julkaisut/workingpapers/2007/ mwp050.htm. November 2011.
* Saidi, T.A. & Tshipala-Ramatshimbila, T.V., 2006. Ecology and management of a remnant Brachystegia spiciformis (miombo) woodland in north eastern Soutpansberg, Limpopo povince. South African Geographical Journal 88(2): 205–212.
* Topps, J.H., 1997. Nutritive value of indigenous browse in Africa in relation to the needs of wild ungulates. Animal Feed Science and Technology 69(1–3): 143–154.
== Author(s) ==
*  L.P.A. Oyen  , PROTA Network Office Europe, Wageningen University, P.O. Box 341, 6700 AH Wageningen, Netherlands *  D. Louppe
* D. Louppe, CIRAD, Département Environnements et Sociétés, Cirad es-dir, Campus international de Baillarguet, TA C 105 / D (Bât. C, Bur. 113), 34398 Montpellier Cédex 5, France
== Correct citation of this article ==
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[[Category:PROTA prov]][[Category:Timbers (PROTA)]]
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