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Copaifera religiosa (PROTA)

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<big>''[[Copaifera religiosa]]'' J.Léonard</big>
 
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{{DISPLAYTITLE:''Copaifera religiosa'' (PROTA)}}
 
:Protologue: Bull. Jard. Bot. Etat 19: 398 (1949).
:Family: Caesalpiniaceae (Leguminosae - Caesalpinioideae)
 
== Synonyms ==
 
 
 
== Vernacular names ==
 
 
== Origin and geographic distribution ==
Bark and roots contain tannin. A dichloromethane bark extract has shown in-vitro antiplasmodial activity against ''Plasmodium falciparum'', while the methanol extract was inactive. The dichloromethane bark extract also showed considerable in-vitro cytotoxicity.
== Botany Description ==
Large tree up to 45 m tall; bole branchless for up to 30 m, straight and cylindrical, up to 200 cm in diameter, thickened at the base, without buttresses; bark surface smooth or finely scaly, orange to reddish brown, inner bark fibrous, brittle, pale yellow, aromatic; crown umbrella-shaped; twigs short-hairy but soon becoming glabrous. Leaves alternate, pinnately compound with 10–16 leaflets; stipules minute, early caducous; petiole and rachis 10–20 cm long, grooved above; petiolules 2–5 mm long, twisted; leaflets alternate, oblong to elliptical or obovate, 2.5–6.5 cm × 1.5–3.5 cm, base slightly asymmetrical and rounded or cuneate, apex rounded to notched, leathery, nearly glabrous, with translucent dots, pinnately veined with c. 20 pairs of lateral veins. Inflorescence an axillary raceme up to 19 cm long, with spike-like branches 2(–6) cm long, densely hairy. Flowers bisexual, zygomorphic, white, nearly sessile, with 2 small, caducous bracteoles; sepals 4, ovate-lanceolate, c. 4 mm × 2–3 mm, 1 broader than other 3, densely hairy; petals absent or sometimes 4 present and then 1–2 mm long; stamens 8–10, free, c. 5 mm long; ovary superior, sessile, hairy at base and sides, 1-celled, style c. 3 mm long. Fruit an ellipsoid to nearly globose pod 3–5 cm × 2–3.5 cm, slightly flattened, leathery, rough, glabrous to hairy, blackish, somewhat shiny, covered with small resin droplets, dehiscent with 2 valves, 1-seeded. Seeds oblong, with a red aril.
 
== Other botanical information ==
''Copaifera'' comprises about 40 species, of which about 35 occur in tropical America, 4 in Africa and 1 in tropical Asia. ''Copaifera'' seems closely related to ''Baikiaea'', ''Detarium'', ''Sindora'' and ''Tessmannia''.
 
== Description ==
 
 
== Ecology ==
* ATIBT (Association Technique Internationale des Bois Tropicaux), 1986. Tropical timber atlas: Part 1 – Africa. ATIBT, Paris, France. 208 pp.
* Aubréville, A., 1970. Légumineuses - Césalpinioidées (Leguminosae - Caesalpinioideae). Flore du Cameroun. Volume 9. Muséum National d’Histoire Naturelle, Paris, France. 339 pp.
* Bolza, E. & Keating, W.G., 1972. African timbers: the properties, uses and characteristics of 700 species. Division of Building Research, CSIRO, Melbourne, Australia. 710 pp.
* Lekana-Douki, J.-B., Oyegue Liabagui, S.L., Bongui, J.B., Zatra, R., Lebibi, J. & Toure-Ndouo, F.S., 2011. In vitro antiplasmodial activity of crude extracts of Tetrapleura tetraptera and Copaifera religiosa. BMC Research Notes 2011, 4: 506. 5 pp.
* Vivien, J. & Faure, J.J., 1985. Arbres des forêts denses d’Afrique Centrale. Agence de Coopération Culturelle et Technique, Paris, France. 565 pp.
== Other references ==
* ATIBT (Association Technique Internationale des Bois Tropicaux), 2007. Statistiques. La lettre de l’ATIBT 26: 38–52.
* Costa, J.A.S., 2009. A new combination in the genus Copaifera (Leguminosae). Neodiversity 4: 14–15.
* de Saint-Aubin, G., 1963. La forêt du Gabon. Publication No 21 du Centre Technique Forestier Tropical, Nogent-sur-Marne, France. 208 pp.
* Koumba Madingou, N.O., Souza, A., Lamidi, M., Mengome, L.E., Eyele Mve Mba, C., Bading Bayissi, Mavoungou, J. & Traore, A.S., 2012. Study of medicinal plants used in the management of cardiovascular diseases at Libreville (Gabon): an ethnopharmacological approach. International Journal of Pharmaceutical Sciences and Research 3(1): 111–119.
* Neuwinger, H.D., 2000. African traditional medicine: a dictionary of plant use and applications. Medpharm Scientific, Stuttgart, Germany. 589 pp.
* Normand, D., 1950. Note sur les bois de Guibourtia arnoldiana et de Copaifera religiosa du Mayumbe. Bulletin du Jardin botanique de l'Etat a Bruxelles 20(1): 19-30.
* Raponda-Walker, A. & Sillans, R., 1961. Les plantes utiles du Gabon. Paul Lechevalier, Paris, France. 614 pp.
* Sallenave, P., 1955. Propriétés physiques et mécaniques des bois tropicaux de l’Union française. Centre Technique Forestier Tropical, Nogent-sur-Marne, France. 129 pp.  * Volavka, Z., 1998. Crown and ritual: the royal insignia of Ngoyo. University of Toronto Press, Toronto, Canada. 411 pp.  * Wilczek, R., Léonard, J., Hauman, L., Hoyle, A.C., Steyaert, R., Gilbert, G. & Boutique, R., 1952. Caesalpiniaceae. In: Robyns, W., Staner, P., Demaret, F., Germain, R., Gilbert, G., Hauman, L., Homès, M., Jurion, F., Lebrun, J., Vanden Abeele, M. & Boutique, R. (Editors). Flore du Congo belge et du Ruanda-Urundi. Spermatophytes. Volume 3. Institut National pour l’Étude Agronomique du Congo belge, Brussels, Belgium. pp. 234–554. == Sources of illustration ==
* Volavka, Z., 1998. Crown and ritual: the royal insignia of Ngoyo. University of Toronto Press, Toronto, Canada. 411 pp.
* Wilczek, R., Léonard, J., Hauman, L., Hoyle, A.C., Steyaert, R., Gilbert, G. & Boutique, R., 1952. Caesalpiniaceae. In: Robyns, W., Staner, P., Demaret, F., Germain, R., Gilbert, G., Hauman, L., Homès, M., Jurion, F., Lebrun, J., Vanden Abeele, M. & Boutique, R. (Editors). Flore du Congo belge et du Ruanda-Urundi. Spermatophytes. Volume 3. Institut National pour l’Étude Agronomique du Congo belge, Brussels, Belgium. pp. 234–554.
== Author(s) ==
*  M. Brink  , PROTA Network Office Europe, Wageningen University, P.O. Box 341, 6700 AH Wageningen, Netherlands
== Correct citation of this article ==
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[[Category:PROTA prov]][[Category:Timbers (PROTA)]]
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